It's kind of cool how he finds bottom end without an actual base. Along with using keys, his combination of fuzz with the whammy pedal creates a super cool bass replacement that you don't hear in other music.
@Jangles exactly you don't notice the absence of bass because the low end is still so prominent
His range makes it work. I don’t think of it here and never felt it was missing from Sleater-Kinney either.